Hybridization and introgression have long complicated relationships within Adenophora and its relatives, challenging generic delimitation. Leveraging deep genome skimming data, Lin et al. (p. 576) generated a large dataset to untangle this reticulate history. Phylogenomic analyses revealed extensive gene tree heterogeneity, with incomplete lineage sorting playing a minimal role. Instead, hybridization and introgression primarily drove early diversification. Integrating phylogenomic, morphological, and geographic evidence, they propose a revised generic framework: expanding Adenophora to include Campanula delavayi and the Korean Peninsula endemic Hanabusaya, reinstating Hyssaria as a distinct Central Asian genus, and establishing two new genera, Boreoasia and Rosomala. The cover shows inflorescence of Adenophora khasiana (云南沙参).

(Image Credit: Yang Niu)